exceeding the limits of physiology, we may treat of their adjusted combinations, so long as these are regarded as parts of the vital consensus. If we observe how the lungs aërate the blood which the heart sends to them; how heart and lungs together supply aërated blood to the stomach, and so enable it to do its work; how these co-operate with sundry secreting and excreting glands to further digestion and to remove waste matter; and how all of them join to keep the brain in a fit condition for carrying on those actions which indirectly conduce to maintenance of the life at large; we are dealing with functions. Even when considering how parts that act directly on the environment – legs, arms, wings – perform their duties, we are still concerned with functions in that aspect of them constituting physiology, so long as we restrict our attention to internal processes, and to internal combinations of them.
But we enter on the subject of conduct when we begin to study such combinations among the actions of sensory and motor organs as are externally manifested. Suppose that instead of observing those contractions of muscles by which the optic axes are converged and the foci of the eyes adjusted (which is a portion of physiology), and that instead of observing the co-operation of other nerves, muscles and bones, by which a hand is moved to a particular place and the fingers closed (which is also a portion of physiology), we observe a weapon being seized by a hand under guidance of the eyes. We now pass from the thought of combined internal functions to the thought of combined external motions. Doubtless, if we could trace the cerebral processes which accompany these, we should find an inner physiological co-ordination corresponding with the outer co-ordination of actions. But this admission is consistent with the assertion, that when we ignore the internal combination and attend only to the external combination, we pass from a portion of physiology to a portion of conduct. For though it may be objected that the external combination instanced is too simple to be rightly included under the name conduct, yet a moment's thought shows that it is joined with what we call conduct by insensible gradations. Suppose the weapon seized is used to ward off a blow. Suppose a counter-blow is given. Suppose the aggressor runs and is chased. Suppose there comes a struggle and a handing him over to the police. Suppose there follow the many and varied acts constituting a prosecution. Obviously the initial adjustment of an act to an end, inseparable from the rest, must be included with them under the same general head; and obviously from this initial simple adjustment, having intrinsically no moral character, we pass by degrees to the most complex adjustments and to those on which moral judgments are passed.
Hence, excluding all internal co-ordinations, our subject here is the aggregate of all external co-ordinations; and this aggregate includes not only the simplest as well as the most complex performed by human beings, but also those performed by all inferior beings considered as less or more evolved.
§ 4. Already the question: What constitutes advance in the evolution of conduct, as we trace it up from the lowest types of living creatures to the highest? has been answered by implication. A few examples will now bring the answer into conspicuous relief.
We saw that conduct is distinguished from the totality of actions by excluding purposeless actions; but during evolution this distinction arises by degrees. In the very lowest creatures most of the movements from moment to moment made, have not more recognizable aims than have the struggles of an epileptic. An infusorium swims randomly about, determined in its course not by a perceived object to be pursued or escaped, but, apparently, by varying stimuli in its medium; and its acts, unadjusted in any appreciable way to ends, lead it now into contact with some nutritive substance which it absorbs, and now into the neighborhood of some creature by which it is swallowed and digested. Lacking those developed senses and motor powers which higher animals possess, ninety-nine in the hundred of these minute animals, severally living for but a few hours, disappear either by innutrition or by destruction. The conduct is constituted of actions so little adjusted to ends, that life continues only as long as the accidents of the environment are favorable. But when, among aquatic creatures, we observe one which, though still low in type, is much higher than the infusorium – say a rotifer – we see how, along with larger size, more developed structures, and greater power of combining functions, there goes an advance in conduct. We see how by its whirling cilia it sucks in as food these small animals moving around; how by its prehensile tail it fixes itself to some fit object; how by withdrawing its outer organs and contracting its body, it preserves itself from this or that injury from time to time threatened; and how thus, by better adjusting its own actions, it becomes less dependent on the actions going on around, and so preserves itself for a longer period.
A superior sub-kingdom, as the Mollusca, still better exemplifies this contrast. When we compare a low mollusc, such as a floating ascidian, with a high mollusc, such as a cephalopod, we are again shown that greater organic evolution is accompanied by more evolved conduct. At the mercy of every marine creature large enough to swallow it, and drifted about by currents which may chance to keep it at sea, or may chance to leave it fatally stranded, the ascidian displays but little adjustment of acts to ends in comparison with the cephalopod; which, now crawling over the beach, now exploring the rocky crevices, now swimming through the open water, now darting after a fish, now hiding itself from some larger animal in a cloud of ink, and using its suckered arms at one time for anchoring itself and at another for holding fast its prey; selects and combines and proportions its movements from minute to minute, so as to evade dangers which threaten, while utilizing chances of food which offer: so showing us varied activities which, in achieving special ends, achieve the general end of securing continuance of the activities.
Among vertebrate animals we similarly trace up, along with advance in structures and functions, this advance in conduct. A fish roaming about at hazard in search of something to eat, able to detect it by smell or sight only within short distances, and now and again rushing away in alarm on the approach of a bigger fish, makes adjustments of acts to ends that are relatively few and simple in their kinds; and shows us, as a consequence, how small is the average duration of life. So few survive to maturity that, to make up for destruction of unhatched young and small fry and half-grown individuals, a million ova have to be spawned by a cod-fish that two may reach the spawning age. Conversely, by a highly-evolved mammal, such as an elephant, those general actions performed in common with the fish are far better adjusted to their ends. By sight as well, probably, as by odor, it detects food at relatively great distances; and when, at intervals, there arises a need for escape, relatively great speed is attained. But the chief difference arises from the addition of new sets of adjustments. We have combined actions which facilitate nutrition – the breaking off of succulent and fruit-bearing branches, the selecting of edible growths throughout a comparatively wide reach; and, in case of danger, safety can be achieved not by flight only, but, if necessary, by defence or attack: bringing into combined use tusks, trunk and ponderous feet. Further, we see various subsidiary acts adjusted to subsidiary ends – now the going into a river for coolness, and using the trunk as a means of projecting water over the body; now the employment of a bough for sweeping away flies from the back; now the making of signal sounds to alarm the herd, and adapting the actions to such sounds when made by others. Evidently, the effect of this more highly-evolved conduct is to secure the balance of the organic actions throughout far longer periods.
And now, on studying the doings of the highest of mammals, mankind, we not only find that the adjustments of acts to ends are both more numerous and better than among lower mammals, but we find the same thing on comparing the doings of higher races of men with those of lower races. If we take any one of the major ends achieved, we see greater completeness of achievement by civilized than by savage; and we also see an achievement of relatively numerous minor ends subserving major ends. Is it in nutrition? The food is obtained more regularly in response to appetite; it is far higher in quality; it is free from dirt; it is greater in variety; it is better prepared. Is it in warmth? The characters of the fabrics and forms of the articles used for clothing, and the adaptations of them to requirements from day to day and hour to hour, are much superior. Is it in dwellings? Between the shelter of boughs and grass which the lowest savage builds, and the mansion of the civilized man, the contrast in aspect is not more extreme than is the contrast in number and efficiency of the adjustments of acts to ends betrayed in their respective constructions. And when with the ordinary activities of the savage we compare the ordinary civilized activities
