all periods.
After the deposit of the Old Red Sandstone, the internal fire appears to have enjoyed a lull of its energy, if not a complete cessation, until the Coal Measures were complete. Then the long tranquility was again broken, and concussions so extensive and violent ensued, that hardly a single square mile of country can anywhere be found which is not full of fractured and contorted strata, the record of subterranean movements, which mostly occurred between the Carboniferous and the Premian deposits.
The effects of these convulsions were manifest in the changed relations of land and sea, existing continents and islands being dislocated, severed, and swallowed up, while others were elevated from the depths of the previous ocean.
It was from the wave-worn materials thus obtained from pre-existing strata, that the New Red Sandstone was consolidated. It consists chiefly of sand and mud, with few organic remains; and the hiatus thus found, in animals and vegetables, seems to be almost a complete one between the organisms of the preceding and the succeeding periods.
The most interesting traces of the earth's tenants during the New Red formation, consist of foot-tracks impressed by the progress of animals along the yielding mud between the ranges of high and low tide. They afford a remarkable example (not, I think, sufficiently dwelt on) of the extreme rapidity with which deposits were consolidated; since the tracks must have been made, and the material consolidated, during the few hours, at most, that intervened between the recess and the reflux of the tide; since, if the mud had not so soon become solid, the flow of the sea would have instantly obliterated such marks, as it does now on our shores.
The principal animal, whose foot-prints have been identified, was an enormous Frog (Labyrinthodon), as big as a hippopotamus, but apparently allied, in its serried teeth, and in the bony plates with which it was covered, to the Crocodiles, which were its associates.
It is curious that marks in the same material have chronicled the serpentine trail of a Sea-worm, the scratchings of a Crab, the ripple of the wavelets, and even the drops of a passing shower; the last revealing, by their margins, the direction of the wind by which the slanting rain was driven.
If the Triassic formations display but little evidence of organic existence, the lack is supplied by the abundance of such records, which is contained in the Oolitic system, and specially in its lowest component, – the Lias. Animals now existed in profusion, but of species which were for the most part peculiar. The coral-making Polypes existed not (or very rarely) in the seas of that age, but lime was secreted by an unusual number of Crinoid Echinoderms, which seem to have fringed the rocks and floating pieces of timber, much as Barnacles do now.
Among the Mollusca now began to appear the inhabitants of those very elegant shells, the Ammonites, allied to the Nautilus of our Southern seas, which may be considered as the lingering representative of those swarms of shelled Cephalopoda. They were accompanied by their near relations, the Belemnites, more resembling a Cuttle, with a long internal, pointed shell.
Fishes, chiefly belonging to a curiously armed tribe of Sharks, together with some enclosed in bony-mail like pavement, were present in the shallows, where the Lias was probably deposited.
But the most characteristic animals were great marine Reptiles, of strange and uncouth forms, to which the present world presents us no known analogy. One of these was the Ichthyosaurus, which closely resembled a porpoise in form, but thirty or forty feet in length, with a vertical fish-like tail, and two pairs of paddles; a mouth set with stout crocodilian teeth, and enormous eyes. Another form was that of the Plesiosaurus, scarcely less in size than its fellow, which in the outline of its body it resembled: it was distinguished, however, by an extraordinary length of neck, slender and swan-like, consisting of thirty or forty vertebræ.
It adds to the interest of these great marine Reptiles, that around their fossil skeletons are preserved pellets of excrement (known as Coprolites) containing fragments of bone, teeth, and scales of fishes, which clearly reveal the nature of their food. In some instances, the stomach and intestines of these great carnivorous creatures, filled with half-digested food, have left indubitable traces of their presence in situ.
Again, the geography of the Globe changed. New lands arose from the sea, and old lands partially or wholly sank. The German Ocean, and part of Western Europe, of our maps, were a great group of islands. The Oolitic formation was deposited. The general character of the organization of this period differed little from that of the Lias. New forms of plants, such as Cycadeæ, were abundant, with, considerable numbers of Corals, Encrinites, Sea-urchins and Mollusks. Macrurous Crustacea, much like those of our times (but essentially different in species), inhabited the sea, and some Beetles and Flies represented the Insects of the land. The Fishes and Marine Reptiles were pretty much the same with those of the Lias, though they received some important additions.
It is, however, among the terrestrial Vertebrata that we must look for the characteristic organisms of this age. And these are, still, Reptiles. The huge Megalosaurus, with a body as big as an elephant's, stood high on his legs, and stretched open a pair of gaping jaws, set with jagged teeth. The Pterodactyles flew about, – carnivorous lizards, with the body and wings of bats,33 except that the membrane was stretched upon the enormously developed little finger; – creatures, perhaps, the most unlike to anything familiar to us, of all fossil forms. And, in the marshy margins of the great river valley which formed the Wealden of our South-eastern districts, the giant Iguanodon, and his fellow, the Hylæosaurus, waged their peaceful warfare on the succulent plants that became their unresisting prey.
The circle of animal life was completed in this epoch, thus far, that every class was represented by some one or more of its constituent species. No fossil skeletons of Birds have, indeed, been found so low as the Oolite, but numerous foot-prints of some of the Grallatores are found in a sandstone of this period; and in the Stonesfield slate, which is contemporary with it, a genus of Mammalia has been discovered, – a small Marsupial, allied to the Opossums of America.
The duration of the Oolitic period must have been considerable. "The lias sea-bottom was succeeded first by a sandy, and then by a calcareous deposit, and the animals were modified accordingly." The deposit of carbonate of lime, which took place under circumstances that caused it to attract around its nodules the organic particles, whence the name oolite (egg-stone) is derived, was not continuous, but repeated at intervals. The shells of Mollusks were developed in great abundance, and accumulations of these formed thick bands, which consolidated into layers of shell-limestone. Three hundred feet of strata, largely composed of organic remains, were formed before the clay was deposited which made the Stonesfield and contemporaneous slates.
Once more the dry land sank, probably by slow successive subsidences, and the sea flowed many fathoms deep above the great European archipelago. And upon its quiet bottom settled down, first a few sandy and clayey beds, and then the great layer of the Chalk.
Creatures of very minute size and low grades of organization were now playing a very important part. A large portion of the lime that was deposited, in the form of a pure carbonate, was doubtless supplied by the Coral structures, which, were exceedingly numerous; the polypidoms being gnawed down by strong-jawed fishes that fed upon the Zoophytes. Foraminifera also were abundant, and contributed to the supply.
Nodules of flint exist in the Chalk, sometimes scattered, sometimes arranged in bands. Two sources are indicated for this substance. One is Sponge, the most common kinds of which are composed of skeletons of siliceous spicula; and these can be discerned with the microscope in the interior of the chalk-flints. But millions upon millions of Infusoria swam through the waters, and many of these were encased in siliceous loricæ, while the rocks and sea-weeds were fringed with as incalculably numerous examples of siliceous Diatomaceæ, whose elegant forms are recognisable without difficulty throughout the Chalk. The inconceivable abundance of these forms may
Mr. Newman suggests that they were "marsupial bats" (Zoologist, p. 129). I have adopted his attitudes, but have not ventured to give them mammalian ears.
