and subsidences of portions of the Earth's crust, bringing about the present irregular distribution of land and sea, have entailed modifications of climate beyond those dependent on latitude; while a yet further series of such modifications have been produced by increasing differences of elevation in the land, which have in sundry places brought arctic, temperate, and tropical climates to within a few miles of one another. And the general outcome of these changes is, that not only has every extensive region its own meteorologic conditions, but that every locality in each region differs more or less from others in those conditions; as in its structure, its contour, its soil. Thus, between our existing Earth, the phenomena of whose crust neither geographers, geologists, mineralogists, nor meteorologists have yet enumerated, and the molten globe out of which it was evolved, the contrast in heterogeneity is extreme.
When from the Earth itself we turn to the plants and animals which have lived, or still live, upon its surface, we find ourselves in some difficulty from lack of facts. That every existing organism has been developed out of the simple into the complex, is indeed the first established truth of all; and that every organism which existed in past times was similarly developed, is an inference no physiologist will hesitate to draw. But when we pass from individual forms of life to Life in general, and inquire whether the same law is seen in the ensemble of its manifestations, – whether modern plants and animals are of more heterogeneous structure than ancient ones, and whether the Earth's present Flora and Fauna are more heterogeneous than the Flora and Fauna of the past, – we find the evidence so fragmentary, that every conclusion is open to dispute. Three-fifths of the Earth's surface being covered by water; a great part of the exposed land being inaccessible to, or untravelled by, the geologist; the greater part of the remainder having been scarcely more than glanced at; and even the most familiar portions, as England, having been so imperfectly explored that a new series of strata has been added within these four years, – it is impossible for us to say with certainty what creatures have, and what have not, existed at any particular period. Considering the perishable nature of many of the lower organic forms, the metamorphosis of numerous sedimentary strata, and the great gaps occurring among the rest, we shall see further reason for distrusting our deductions. On the one hand, the repeated discovery of vertebrate remains in strata previously supposed to contain none, – of reptiles where only fish were thought to exist, – of mammals where it was believed there were no creatures higher than reptiles, – renders it daily more manifest how small is the value of negative evidence. On the other hand, the worthlessness of the assumption that we have discovered the earliest, or anything like the earliest, organic remains, is becoming equally clear. That the oldest known sedimentary rocks have been greatly changed by igneous action, and that still older ones have been totally transformed by it, is becoming undeniable. And the fact that sedimentary strata earlier than any we know, have been melted up, being admitted, it must also be admitted that we cannot say how far back in time this destruction of sedimentary strata has been going on. Thus the title Palæozoic, as applied to the earliest known fossiliferous strata, involves a petitio principii; and, for aught we know to the contrary, only the last few chapters of the Earth's biological history may have come down to us. On neither side, therefore, is the evidence conclusive. Nevertheless we cannot but think that, scanty as they are, the facts, taken altogether, tend to show both that the more heterogeneous organisms have been evolved in the later geologic periods, and that Life in general has been more heterogeneously manifested as time has advanced. Let us cite, in illustration, the one case of the Vertebrata. The earliest known vertebrate remains are those of Fishes; and Fishes are the most homogeneous of the vertebrata. Later and more heterogeneous are Reptiles. Later still, and more heterogeneous still, are Birds and Mammals. If it be said that the Palæozoic deposits, not being estuary deposits, are not likely to contain the remains of terrestrial vertebrata, which may nevertheless have existed at that era, we reply that we are merely pointing to the leading facts, such as they are. But to avoid any such criticism, let us take the mammalian subdivision only. The earliest known remains of mammals are those of small marsupials, which are the lowest of the mammalian type; while, conversely, the highest of the mammalian type – Man – is the most recent. The evidence that the vertebrate fauna, as a whole, has become more heterogeneous, is considerably stronger. To the argument that the vertebrate fauna of the Palæozoic period, consisting, so far as we know, entirely of Fishes, was less heterogeneous than the modern vertebrate fauna, which includes Reptiles, Birds, and Mammals, of multitudinous genera, it may be replied, as before, that estuary deposits of the Palæozoic period, could we find them, might contain other orders of vertebrata. But no such reply can be made to the argument that whereas the marine vertebrata of the Palæozoic period consisted entirely of cartilaginous fishes, the marine vertebrata of later periods include numerous genera of osseous fishes; and that, therefore, the later marine vertebrate faunas are more heterogeneous than the oldest known one. Nor, again, can any such reply be made to the fact that there are far more numerous orders and genera of mammalian remains in the tertiary formations than in the secondary formations. Did we wish merely to make out the best case, we might dwell upon the opinion of Dr. Carpenter, who says that "the general facts of Palæontology appear to sanction the belief, that the same plan may be traced out in what may be called the general life of the globe, as in the individual life of every one of the forms of organized being which now people it." Or we might quote, as decisive, the judgment of Professor Owen, who holds that the earlier examples of each group of creatures severally departed less widely from archetypal generality than the later examples – were severally less unlike the fundamental form common to the group as a whole; and thus constituted a less heterogeneous group of creatures. But in deference to an authority for whom we have the highest respect, who considers that the evidence at present obtained does not justify a verdict either way, we are content to leave the question open.2
Whether an advance from the homogeneous to the heterogeneous is or is not displayed in the biological history of the globe, it is clearly enough displayed in the progress of the latest and most heterogeneous creature – Man. It is true alike that, during the period in which the Earth has been peopled, the human organism has grown more heterogeneous among the civilized divisions of the species; and that the species, as a whole, has been growing more heterogeneous in virtue of the multiplication of races and the differentiation of these races from each other. In proof of the first of these positions, we may cite the fact that, in the relative development of the limbs, the civilized man departs more widely from the general type of the placental mammalia than do the lower human races. While often possessing well-developed body and arms, the Australian has very small legs: thus reminding us of the chimpanzee and the gorilla, which present no great contrasts in size between the hind and fore limbs. But in the European, the greater length and massiveness of the legs have become marked – the fore and hind limbs are more heterogeneous. Again, the greater ratio which the cranial bones bear to the facial bones illustrates the same truth. Among the vertebrata in general, progress is marked by an increasing heterogeneity in the vertebral column, and more especially in the segments constituting the skull: the higher forms being distinguished by the relatively larger size of the bones which cover the brain, and the relatively smaller size of those which form the jaws, &c. Now this characteristic, which is stronger in Man than in any other creature, is stronger in the European than in the savage. Moreover, judging from the greater extent and variety of faculty he exhibits, we may infer that the civilized man has also a more complex or heterogeneous nervous system than the uncivilized man: and, indeed, the fact is in part visible in the increased ratio which his cerebrum bears to the subjacent ganglia, as well as in the wider departure from symmetry in its convolutions. If further elucidation be needed, we may find it in every nursery. The infant European has sundry marked points of resemblance to the lower human races; as in the flatness of the alæ of the nose, the depression of its bridge, the divergence and forward opening of the nostrils, the form of the lips, the absence of a frontal sinus, the width between the eyes, the smallness of the legs. Now, as the developmental process by which these traits are turned into those of the adult European, is a continuation of that change from the homogeneous to the heterogeneous displayed during the previous evolution of the embryo, which every anatomist will admit; it follows that the parallel developmental process by which the like traits of the barbarous races have been turned into those of the civilized races, has also been a continuation of the change from the homogeneous
Since this was written (in 1857) the advance of paleontological discovery, especially in America, has shown conclusively, in respect of certain groups of vertebrates, that higher types have arisen by modifications of lower; so that, in common with others, Prof. Huxley, to whom the above allusion is made, now admits, or rather asserts, biological progression, and, by implication, that there have arisen more heterogeneous organic forms and a more heterogeneous assemblage of organic forms.
