this statement is only an adumbration. The order of Nature is habitually more complex than our generalizations represent it as being – refuses to be fully expressed in simple formulæ; and we are obliged to limit them by various qualifications. It is thus here. Since von Baer's day the careful observations of numerous observers have shown his allegation to be but approximately true. Hereafter, when discussing the embryological evidence of Evolution, the causes of deviations will be discussed. For the present it suffices to recognize as unquestionable the fact that whereas the germs of organisms are extremely similar, they gradually diverge widely, in modes now regular and now irregular, until in place of a multitude of forms practically alike we finally have a multitude of forms most of which are extremely unlike. Thus, in conformity with the law of evolution, not only do the parts of each organism advance from indefinite homogeneity to definite heterogeneity, but the assemblage of all organisms does the same: a truth already indicated in First Principles.
§ 53. This comparison between the course of development, in any creature, and the course of development in all other creatures – this arrival at the conclusion that the course of development in each, at first the same as in all others, becomes stage by stage differentiated from the courses in all others, brings us within view of an allied conclusion. If we contemplate the successive stages passed through by any higher organism, and observe the relation between it and its environment at each of these stages; we shall see that this relation is modified in a way analogous to that in which the relation between the organism and its environment is modified, as we advance from the lowest to the highest grades. Along with the progressing differentiation of each organism from others, we find a progressing differentiation of it from its environment; like that progressing differentiation from the environment which we meet with in the ascending forms of life. Let us first glance at the way in which the ascending forms of life exhibit this progressing differentiation from the environment.
In the first place, it is illustrated in structure. Advance from the homogeneous to the heterogeneous, itself involves an increasing distinction from the inorganic world. Passing over the Protozoa, of which the simplest probably disappeared during the earliest stages of organic evolution, and limiting our comparison to the Metazoa, we see that low types of these, as the Cœlenterata, are relatively simple in their organization; and the ascent to organisms of greater and greater complexity of structure, is an ascent to organisms which are in that respect more strongly contrasted with the structureless environment. In form, again, we see the same truth. An ordinary characteristic of inorganic matter is its indefiniteness of form; and this is also a characteristic of the lower organisms, as compared with the higher. Speaking generally, plants are less definite than animals, both in shape and size – admit of greater modifications from variations of position and nutrition. Among animals, the simplest Rhizopods may almost be called amorphous: the form is never specific, and is constantly changing. Of the organisms resulting from the aggregation of such creatures, we see that while some, as the Foraminifera, assume a certain definiteness of form, in their shells at least, others, as the Sponges, are very irregular. The Zoophytes and the Polyzoa are compound organisms, most of which have a mode of growth not more determinate than that of plants. But among the higher animals, we find not only that the mature shape of each species is very definite, but that the individuals of each species differ little in size. A parallel increase of contrast is seen in chemical composition. With but few exceptions, and those only partial ones, the lowest animal and vegetal forms are inhabitants of the water; and water is almost their sole constituent. Desiccated Protophyta and Protozoa shrink into mere dust; and among the Acalephes we find but a few grains of solid matter to a pound of water. The higher aquatic plants, in common with the higher aquatic animals, possessing as they do increased tenacity of substance, also contain a greater proportion of the organic elements; further they show us a greater variety of composition in their different parts; and thus in both ways are chemically more unlike their medium. And when we pass to the superior classes of organisms – land-plants and land-animals – we see that, chemically considered, they have little in common either with the earth on which they stand or the air which surrounds them. In specific gravity too, we may note a like truth. The simplest forms, in common with the spores and gemmules of higher ones, are as nearly as may be of the same specific gravity as the water in which they float; and though it cannot be said that among aquatic creatures, superior specific gravity is a standard of general superiority, yet we may fairly say that the higher orders of them, when divested of the appliances by which their specific gravity is regulated, differ more from water in their relative weights than do the lowest. In terrestrial organisms, the contrast becomes marked. Trees and plants, in common with insects, reptiles, mammals, birds, are all of a specific gravity considerably less than that of the earth and immensely greater than that of the air. Yet further, we see the law fulfilled in respect of temperature. Plants generate but extremely small quantities of heat, which are to be detected only by delicate experiments; and practically they may be considered as having the same temperature as their environment. The temperature of aquatic animals is very little above that of the surrounding water: that of the invertebrata being mostly less than a degree above it, and that of fishes not exceeding it by more than two or three degrees; save in the case of some large red-blooded fishes, as the tunny, which exceed it in temperature by nearly ten degrees. Among insects the range is from two to ten degrees above that of the air: the excess varying according to their activity. The heat of reptiles is from four to fifteen degrees more than the heat of their medium. While mammals and birds maintain a heat which continues almost unaffected by external variations, and is often greater than that of the air by seventy, eighty, ninety, and even a hundred degrees. Once more, in greater self-mobility a progressive differentiation is traceable. The chief characteristic by which we distinguish dead matter is its inertness: some form of independent motion is our most familiar proof of life. Passing over the indefinite border-land between the animal and vegetal kingdoms, we may roughly class plants as organisms which, while they exhibit that kind of motion implied in growth, are not only devoid of locomotive power, but with some unimportant exceptions are devoid of the power of moving their parts in relation to each other; and thus are less differentiated from the inorganic world than animals. Though in those microscopic Protophyta and Protozoa inhabiting the water we see locomotion produced by ciliary action; yet this locomotion, while rapid relatively to the sizes of their bodies, is absolutely slow. Of the Cœlenterata a great part are either permanently rooted or habitually stationary; and so have scarcely any self-mobility but that implied in the relative movements of parts; while the rest, of which the common jelly-fish serves as a sample, have mostly but little ability to move themselves through the water. Among the higher aquatic Invertebrata, – cuttlefishes and lobsters, for instance, – there is a very considerable power of locomotion; and the aquatic Vertebrata are, considered as a class, much more active in their movements than the other inhabitants of the water. But it is only when we come to air-breathing creatures that we find the vital characteristics of self-mobility manifested in the highest degree. Flying insects, mammals, birds, travel with velocities far exceeding those attained by any of the lower classes of animals. Thus, on contemplating the various grades of organisms in their ascending order, we find them more and more distinguished from their inanimate media, in structure, in form, in chemical composition, in specific gravity, in temperature, in self-mobility. It is true that this generalization does not hold with complete regularity. Organisms which are in some respects the most strongly contrasted with the environing inorganic world, are in other respects less contrasted than inferior organisms. As a class, mammals are higher than birds; and yet they are of lower temperature and have smaller powers of locomotion. The stationary oyster is of higher organization than the free-swimming medusa; and the cold-blooded and less heterogeneous fish is quicker in its movements than the warm-blooded and more heterogeneous sloth. But the admission that the several aspects under which this increasing contrast shows itself, bear variable ratios to each other, does not conflict with the general truth that as we ascend in the hierarchy of organisms, we meet with not only an increasing differentiation of parts but also an increasing differentiation from the surrounding medium in sundry other physical attributes. It would seem that this trait has some necessary connexion with superior vital manifestations. One of those lowly gelatinous forms, so transparent and colourless as to be with difficulty distinguished from the water it floats in,
