toes, occasionally wanting (as in the Hemipods). Tarsus, when not feathered, generally broadly scutellate. Front toes commonly webbed at base. Claws blunt, little curved. Wings strong, short, and concavo-convex. Rectrices commonly more than twelve. Head small. Plumage usually after-shafted. Carotids double (except Turnicidæ and Megapodidæ). No intrinsic syringeal muscles. Sternum very deeply, generally doubly, notched. Palate schizognathous. Chiefly polygamous. Præcocial and ptilopædic.
G. LIMICOLÆ. Tibiæ bare of feathers for a variable (sometimes very slight) distance above the suffrago. Legs commonly lengthened, sometimes excessively so, and neck usually produced in corresponding ratio. Tarsi scutellate or reticulate. Toes never coherent at base; cleft, or united for a short distance by one or two small movable basal webs (palmate only in Recurvirostra, lobate only in Phalaropodidæ). Hallux always reduced, obviously elevated and free, or wanting; giving a foot of cursorial character. Wings, with few exceptions, lengthened, pointed, and flat; the inner primaries and outer secondaries very short, forming a strong re-entrance on the posterior border of the wing. Tail shorter than the wing, of simple form, and of few feathers, except in certain Snipes. Head globose, sloping rapidly down to the contracted base of the bill, completely feathered (except Philomachus ♂). Gape of bill short and constricted; tip usually obtuse; bill weak and flexible. Rostrum commonly lengthened, and more or less terete and slender; membranous wholly or in great part, without hard cutting edges. Nostrils narrow, placed low down, entirely surrounded with soft skin; nasal fossæ extensive. Palate schizognathous. Sternum usually doubly, sometimes singly, notched. Carotids double. Pterylosis of a particular pattern. Nature præcocial and ptilopædic. Comprising the “Plover-Snipe” group; species of medium and small size, with never extremely compressed or depressed body; more or less aquatic, living on plains and in open places, usually near water, nesting on the ground, where the young run freely at birth.
H. HERODIONES. Tibiæ naked below. Legs and neck much lengthened in corresponding ratio. Toes long, slender, never coherent at base, where cleft, or with movable basal webbing. Hallux (as compared with that of the preceding and following group) lengthened, free, and either perfectly incumbent or but little elevated, with a large claw, giving a foot of insessorial character. Wings commonly obtuse, but broad and ample, without marked re-entrance on posterior border, the intermediate remiges not being much abbreviated. Tail short and few-feathered. Head narrow, conico-elongated, gradually contracting to the large, stout base of the bill; the loral and orbital region, or the whole head, naked. Gape of the bill deeply fissured; tip usually acute; tomia hard and cutting. Bill conico-elongate, always longer than the head, stout and firm. Nostrils small, placed high up, with entirely bony and horny, or only slightly membranous, surroundings. Pterylosis nearly peculiar in the presence, almost throughout the group, of powder-down tracts, rarely found elsewhere; pterylæ very narrow. Palate desmognathous. Carotids double. Altricial. Comprising the Herons, Storks, Ibises, etc. (not Cranes). Species usually of large stature, with compressed body and very long S-bent neck; perching and nesting usually in trees, bushes, or other high places near water; young hatching weak, scarcely feathered, and reared in the nest.
I. ALECTORIDES. 12 Tibiæ naked below. Neck, legs, and feet much as in the last group, but hallux reduced and obviously elevated, with small claw, the resulting foot cursorial (natatorial and lobate in Fulica). Wings and tail commonly as in Herodiones. Head less narrowed and conic than in the last, fully feathered or with extensive baldness (not with definite nakedness of loral and orbital regions). Bill of various shape, usually lengthened and obtuse, never extensively membranous. Rictus moderate. Nostrils lower than in Herodiones. Pterylosis not peculiar. Palate schizognathous. Carotids double. Nature præcocial and ptilopædic. Comprising the Cranes and Rails and their allies; the former agreeing with the Herodiones superficially in stature, etc., but highly diverse in the schizognathous palate, præcocial nature, etc.
J. LAMELLIROSTRES. Feet palmate; tibiæ feathered (except Phœnicopterus). Legs near centre of equilibrium of the body, its axis horizontal in walking; not lengthened except in Phœnicopterus. Knee-joint rarely exserted beyond general skin of the body. Wings moderate, reaching when folded to, but not beyond, the usually short and rounded (exceptionally long and cuneate) tail. Feet tetradactyle (except sometimes in Phœnicopterus); hallux reduced, elevated and free, often independently lobate. Bill lamellate, i. e., furnished along each commissural edge with a regular series of mutually adapted laminæ or tooth-like processes, with which correspond certain laciniate processes of the fleshy tongue, which ends in a horny tip. Bill large, thick, high at base, depressed towards the end, membranous to the broad obtuse tip, which is occupied by a horny “nail” of various shape. Nostrils patent, never tubular; nasal fossæ slight. No gular pouch. Plumage dense, to resist water. Eyes very small. Head high, compressed, with lengthened, sloping frontal region. Palate desmognathous. Reproduction præcocial; young ptilopædic. Eggs numerous. Carotids double. Sternum single-notched. Comprising Flamingoes and all the Anserine birds.
K. STEGANOPODES. Feet totipalmate; hallux lengthened, nearly incumbent, semilateral, completely united with the second toe by a full web. Tibiæ feathered; position of legs with reference to axis of body variable, but generally far posterior; knee-joint not free. Wings and tail variable. Bill of very variable shape, never lamellate, wholly corneous; its tomia often serrate; external nares very small or finally abortive. A prominent naked gular pouch. Tarsi reticulate. Sternum entire or nearly so; furculum confluent with its keel. Carotids double. Palate highly desmognathous. Reproduction altricial; young psilopædic or ptilopædic. Eggs three or fewer.
L. LONGIPENNES. (To most of the characters of the group here given the genus Halodroma is a signal exception, though unquestionably belonging here.) Feet palmate. Tibiæ feathered. Legs at or near centre of equilibrium, affording horizontal position of axis of body in walking. Knee scarcely buried in common integument; tibia sometimes with a long apophysis. Hallux elevated, free, functionless; very small, rudimentary, or wanting. Rostrum of variable shape, usually compressed and straight to the hooked end, sometimes entirely straight and acute, commonly lengthened, always corneous, without serration or true lamellæ. Nostrils of various forms, tubular or simply fissured, never abortive. No gular pouch. Wings very long and pointed, surpassing the base and often the end of the large, well-formed, few-feathered tail. Carotids double. Palate schizognathous. Reproduction altricial; young ptilopædic. Eggs three or fewer. Habit highly volucral.
M. PYGOPODES. Feet palmate or lobate. Tibiæ feathered, often with a long apophysis, always buried in common integument nearly to the heel-joint, necessitating a more or less erect posture of the body on land, where progression is difficult. Hallux small, elevated or wanting; feet lobate or palmate. Bill of indeterminate shape, wholly corneous, never lamellate or serrate, nor with gular pouch. Nostrils not abortive. Wings very short, reaching scarcely or not to the base, never to the tip, of the short, sometimes rudimentary, tail. Palate schizognathous. Carotid usually double, sometimes single (in Podiceps and Mergulus). Nature altricial or præcocial; young ptilopædic. Highly natatorial.
N. SPHENISCI. With general characters of the last group, but distinguished by unique ptilosis and wing-structure, etc. Plumage without apteria, of singularly modified scale-like feathers on most parts; no developed remiges. Wings unfit for flight, insusceptible of perfect flexion or extension, very short, with peculiarly flattened bones and stable articulations. Skeleton non-pneumatic. Many bones, terete in ordinary birds, here flattened. Metatarsal bone flattened transversely, doubly fenestrate. Hallux elevated, lateral, minute, free. No free pollex. Two anconal sesamoids; patella from double centres; tibia without apophysis; a free tarsal ossicle. Sternum with long lateral apophyses. Pelvic connections unstable. Carotids double. Comprising only the Penguins. Confined to the Southern Hemisphere.
Having thus presented and defined an arrangement of the higher groups into which recent Carinate birds are susceptible of division, I next proceed to the consideration of the North American Families of birds which the authors of the present work have provisionally adopted as suitable to the
Groups G., H., and I. are respectively equal to the
