Natural History of the Mammalia of India and Ceylon. Robert Armitage Sterndale
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DESCRIPTION.—Front and sides of the face, outside of fore-limbs, throat and chest, golden yellow; inner side of hind limbs rich red brown, which is also the colour of the hind legs and feet; head dark brown, with golden hairs intermixed; back dark maroon, almost black; upper surface of the tail the same; pale oval patch between shoulders, dark band on each side between it and fore-limbs, passing forward over the ears.
SIZE.—Head and body, 7·10; tail, 8 inches.
There is a little animal allied to the genus Tupaia, which has hitherto been found only in Borneo and Sumatra, but as Sumatran types have been found in Tenasserim, perhaps some day the Ptilocercus Lowii may be discovered there. It has a rather shorter head than the true Banxrings, more like T. Ellioti, but its dentition is nearly the same, as also are its habits. Its chief peculiarity lies in its tail, which is long, slender and naked, like that of a rat for two-thirds of its length, the terminal third being adorned with a broad fringe of hair on each side, like the wings of an arrow or the plumes of a feather. There is an excellent coloured picture of it in the 'Proc. Zool. Society,' vol. of Plates.
I had almost concluded my sketch of the Insectivora without alluding to one most interesting genus, which ought properly to have come between the shrews and the hedgehogs, the Gymnura, which, though common in the Malay countries, has only recently been found in Burmah—a fact of which I was not aware till I saw it included in a paper on Tenasserim mammals by Mr. W. T. Blanford ('Jour. As. Soc. Beng.,' 1878, page 150). Before I refer to his notes I may state that this animal is a sort of link between the Soricidæ and the Erinaceidæ, and De Blainville proposed for it the generic name of Echinosorex, but the one generally adopted is Gymnura, which was the specific name given to it by its discoverer, Sir Stamford Raffles, who described it as a Viverra (V. gymnura); however, Horsfield and Vigors and Lesson, the two former in England and the latter in France, saw that it was not a civet, and, taking the naked tail as a peculiarity, they called the genus Gymnura, and the specimen Rafflesii. There is not much on record regarding the anatomy of the animal, and in what respects it internally resembles the hedgehogs. Outwardly it has the general soricine form, though much larger than the largest shrew. The long tail too is against its resemblance to the hedgehogs, which rests principally on its spiny pelage.
The teeth in some degree resemble Erinaceus, the molars and premolars especially, but the number in all is greater, there being forty-four, or eight more. It would be interesting to know whether the zygomatic arch is perfect and the tibia and fibula united, as in the hedgehogs, or wanting and distinct as in the shrews. I have given a slight sketch in outline of the animal.
HABITAT.—Tenasserim (Sumatra, Borneo); Malacca.
DESCRIPTION.—Long tapering head, with elongated muzzle, short legs, shrew-like body, with a long, round, tapering and scaly rat-like tail, naked, with the exception of a few stiff hairs here and there among the scales. In each jaw on each side three incisors, one canine (those in the upper jaw double-fanged) and seven premolars and molars; feet five-toed, plantigrade, armed with strong claws. Fur of two kinds, fine and soft, with longer and more spiny ones intermixed. The colour varies a good deal, the general tint being greyish-black, with head and neck pale or whitish, and with a broad black patch over the eye. Some have been found almost wholly white, with the black eye-streak and only a portion of the longer hairs black, so that much stress cannot be laid on the colouring; the tail is blackish at the base, whitish and compressed at the tip. Mr. Blanford says: "The small scales covering the tail are indistinctly arranged in rings and sub-imbricate; on the lower surface the scales are convex and distinctly imbricate, the bristles arising from the interstices. Thus the under surface of the tail is very rough, and may probably be of use to the animal in climbing." He also refers to the fact that the claws of his specimen are not retractile, and mentions that in the original description both in Latin and English the retractability of the claws is pointed out as a distinction between Gymnura and Tupaia. In the description given of the Sumatran animal both by Dallas and Cuvier nothing is mentioned about this feature.
SIZE.—A Sumatran specimen: head and body, 14 inches; tail, 12 inches. Mr. Blanford's specimen: head and body, 12 inches; tail, 8·5.
Mr. Blanford was informed by Mr. Davison, who obtained it in Burmah, that the Gymnura is purely nocturnal in its habits, and lives under the roots of trees. It has a peculiar and most offensive smell, resembling decomposed cooked vegetables. The Bulau has not the power of rolling itself up like the hedgehog, nor have the similar forms of insectivores which resemble the hedgehog in some respects, such as the Tenrecs (Centetes), Tendracs (Ericulus), and Sokinahs (Echinops) of Madagascar.
Speaking generally, the whole range of mammals between the Quadrumana and the Rodentia are carnivorous with few exceptions, yet there is one family which, from its muscular development and dentition, is pre-eminently flesh-eating, as Cuvier aptly remarks, "the sanguinary appetite is combined with the force necessary for its gratification." Their forms are agile and muscular; their circulation and respiration rapid. As Professor Kitchen Parker graphically writes: "This group, which comprises all the great beasts of prey, is one of the most compact as well as the most interesting among the mammalia. So many of the animals contained in it have become 'familiar in our mouths as household words,' bearing as they do an important part in fable, in travel, and even in history; so many of them are of such wonderful beauty, so many of such terrible ferocity, that no one can fail to be interested in them, even apart from the fact likely to influence us more in their favour than any other, that the two home pets, which of all others are the commonest and the most interesting, belong to the group. No one who has had a dog friend, no one who has watched the wonderful instance of maternal love afforded by a cat with her kittens, no one who loves riding across country after a fox, no lady with a taste for handsome furs, no boy who has read of lion and tiger hunts and has longed to emulate the doughty deeds of the hunter, can fail to be interested in an assemblage which furnishes animals at once so useful, so beautiful and so destructive. It must not be supposed from the name of this group that all its members are exclusively flesh-eaters, and indeed it will be hardly necessary to warn the reader against falling into this mistake, as there are few people who have never given a dog a biscuit, or a bear a bun. Still both the dog and several kinds of bears prefer flesh-meat when they can get it, but there are some bears which live almost exclusively on fruit, and are, therefore, in strictness not carnivorous at all. The name must, however, be taken as a sort of general title for a certain set of animals which have certain characteristics in common, and which differ from all other animals in particular ways." I would I had more space at my disposal for further quotations from Professor Parker's 'General Remarks on the Land Carnivora,' his style is so graphic.
The dentition of the Carnivora varies according to the exclusiveness of their fleshy diet, and the nature of that diet.
In taking two typical forms I give below sketches from skulls in my possession of the tiger, and the common Indian black bear; the one has trenchant cutting teeth which work up and down, the edges sliding past each other just like a pair of scissors; the other has flat crowned molars adapted for triturating the roots and herbage on which it feeds. A skull of an old bear which I have has molars of which the crowns are worn almost smooth from attrition. In the most carnivorous forms the tubercular molars are almost rudimentary.
The skull exhibits peculiar features for the attachment of the necessary powerful muscles. The bones of the face are short in comparison with the cranial portion of the skull (the reverse of the Herbivores); the strongly built zygomatic arch, the roughened ridges and the broad ascending ramus of the lower jaw, all afford place for the attachment of the immense muscular development. Then the hinge of the jaw is peculiar; it allows of no