to the cell doctrine of Schleiden and Schwann, matured about 1840, which remains, with some modifications, the basis of modern biology. The advance along these lines contributed signally to the advancement of embryology, which reached a startling height in 1829 with the publication of Baer's memorable treatise, in which the development of an ovum is shown to consist in a change from homogeneity to heterogeneity through successive differentiations. But while Baer thus arrived at the very threshold of the law of evolution, he was not in the true sense an evolutionist; he had nothing to say to phylogenetic evolution, or the derivation of the higher forms of life from lower forms through physical descent with modifications. Just so with Cuvier. When he effected his grand classification, he prepared the way most thoroughly for a general theory of evolution, but he always resisted any such inference from his work. He was building better than he knew.
The hesitancy of such men as Cuvier and Baer was no doubt due partly to the apparent absence of any true cause for physical modifications in species, partly to the completeness with which their own great work absorbed their minds. Often in the history of science we witness the spectacle of a brilliant discoverer travelling in triumph along some new path, but stopping just short of the goal which subsequent exploration has revealed. There it stands looming up before his face, but he is blind to its presence through the excess of light which he has already taken in. The intellectual effort already put forth has left no surplus for any further sweep of comprehension, so that further advance requires a fresher mind and a new start with faculties unjaded and unwarped. To discover a great truth usually requires a succession of thinkers. Among the eminent anatomists who in the earlier part of our century were occupied with the classification of animals, there were some who found themselves compelled to believe in phylogenetic evolution, although they could frame no satisfactory theory to account for it. The weight of evidence was already in favour of such evolution, and these men could not fail to see it. Foremost among them was Jean Baptiste Lamarck, whose work was of supreme importance. His views were stated in 1809 in his "Philosophic Zoölogique," and further illustrated in 1815, in his voluminous treatise on invertebrate animals. Lamarck entirely rejected the notion of special creations, and he pointed out some of the important factors in evolution, especially the law that organs and faculties tend to increase with exercise, and to diminish with disuse. His weakest point was the disposition to imagine some inherent and ubiquitous tendency toward evolution, whereas a closer study of nature has taught us that evolution occurs only where there is a concurrence of favourable conditions. Among others who maintained some theory of evolution were the two Geoffroy Saint-Hilaires, father and son, and the two great botanists, Naudin in France and Hooker in England. In 1852 the case of evolution as against special creations was argued by Herbert Spencer with convincing force, and in 1855 appeared "The Principles of Psychology," by the same author, a book which is from beginning to end an elaborate illustration of the process of evolution, and is divided from everything that came before it by a gulf as wide as that which divides the Copernican astronomy from the Ptolemaic.
The followers of Cuvier regarded the methods and results of these evolutionists with strong disapproval. In the excess of such a feeling, they even went so far as to condemn all philosophic thinking on subjects within the scope of natural history as visionary and unscientific. Why seek for any especial significance in the fact that every spider and every lobster is made up of just twenty segments? Is it not enough to know the fact? Children must not ask too many questions. It is the business of science to gather facts, not to seek for hidden implications. Such was the mental attitude into which men of science were quite commonly driven, between 1830 and 1860, by their desire to blink the question of evolution. A feeling grew up that the true glory of a scientific career was to detect for the two hundredth time an asteroid, or to stick a pin through a beetle with a label attached bearing your own latinized name, Browni, or Jonesii, or Robinsoniense. This feeling was especially strong in France, and was not confined to physical science. It was exhibited a few years later in the election of some Swedish or Norwegian naturalist (whose name I forget) to the French Academy of Science instead of Charles Darwin: the former had described some new kind of fly, the latter was only a theorizer! The study of origins in particular was to be frowned upon. In 1863 the Linguistic Society of Paris passed a by-law that no communications bearing upon the origin of language would be received. In the same mood, Sir Henry Maine's treatise on "Ancient Law" was condemned at a leading American university: it was enough for us to know our own laws; those of India might interest British students who might have occasion to go there, but not Americans. Such crude notions, utterly hostile to the spirit of science, were unduly favoured fifty years ago by the persistent unwillingness to submit the phenomena of organic nature to the kind of scientific explanation which facts from all quarters were urging upon us.
During the period from 1830 to 1860, the factor in evolution which had hitherto escaped detection was gradually laid hold of and elaborately studied by Charles Darwin. In the nature of his speculations, and the occasion that called them forth, he was a true disciple of Lyell. The work of that great geologist led directly up to Darwinism. As long as it was supposed that each geologic period was separated from the periods before and after it by Titanic convulsions which revolutionized the face of the globe, it was possible for men to acquiesce in the supposition that these convulsions wrought an abrupt and a wholesale destruction of organic life, and that the lost forms were replaced by an equally abrupt and wholesale supernatural creation of new forms at the beginning of each new period. But, as people ceased to believe in the convulsions, such an explanation began to seem improbable, and it was completely discredited by the fact that many kinds of plants and animals have persisted with little or no change during several successive periods, side by side with other kinds in which there have been extensive variation and extinction.
In connection with this a fact of great significance was elicited. Between the fauna and flora of successive periods in the same geographical region there is apt to be a manifest family likeness, indicating that the later are connected with the earlier through the bonds of physical descent. It was a case of this sort that attracted Darwin's attention in 1835. The plants and animals of the Galapagos Islands are either descended, with specific modifications, from those of the mainland of Ecuador, or else there must have been an enormous number of special creations. The case is one which at a glance presents the notion of special creations in an absurd light. But what could have caused the modification? What was wanted was, to be able to point to some agency, similar to agencies now in operation, and therefore intelligible, which could be proved to be capable of making specific changes in plants and animals. Darwin's solution of the problem was so beautiful, it seems now so natural and inevitable, that we may be in danger of forgetting how complicated and abstruse the problem really was. Starting from the known experiences of breeders of domestic animals and cultivated plants, and duly considering the remarkable and sometimes astonishing changes that are wrought by simple selection, the problem was to detect among the multifarious phenomena of organic nature any agency capable of accomplishing what man thus accomplishes by selection. In detecting the agency of natural selection, working perpetually through the preservation of favoured individuals and races in the struggle for existence, Darwin found the true cause for which men were waiting. With infinite patience and caution, he applied his method of explanation to one group of organic phenomena after another, meeting in every quarter with fresh and often unexpected verification. After more than twenty years, a singular circumstance led him to publish an account of his researches. The same group of facts had set a younger naturalist to work upon the same problem, and a similar process of thought had led to the same solution. Without knowing what Darwin had done, Alfred Russel Wallace made the same discovery, and sent from the East Indies, in 1858, his statement of it to Darwin as to the man whose judgment upon it he should most highly prize. This made publication necessary for Darwin. The vast treasures of theory and example which he had accumulated were given to the world, the notion of special creations was exploded, and the facts of phylogenetic evolution won general acceptance.
Under the influence of this great achievement, men in every department of science began to work in a more philosophical spirit. Naturalists, abandoning the mood of the stamp collectors, saw in every nook and corner some fresh illustration of Darwin's views. One serious obstacle to any general statement of the doctrine of evolution was removed. It was in 1861 that Herbert Spencer began to publish such a general
