Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras. Alfred Russel Wallace
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Migrations and Extinction of Organisms caused by the Glacial Epoch.—Our last glacial epoch was accompanied by at least two considerable submergences and elevations of the land, and there is some reason to think, as we have already explained, that the two classes of phenomena are connected as cause and effect. We can easily see how such repeated submergences and elevations would increase and aggravate the migrations and extinctions that a glacial epoch is calculated to produce. We can therefore hardly fail to be right in attributing the wonderful changes in animal and vegetable life that have occurred in Europe and N. America between the Miocene Period and the present day, in part at least, to the two or more cold epochs that have probably intervened. These changes consist, first, in the extinction of a whole host of the higher animal forms, and secondly, in a complete change of types due to extinction and migration, leading to a much greater difference between the vegetable and animal forms of the eastern and western hemisphere than before existed. Many large and powerful mammalia lived in our own country in Pliocene times and apparently survived a part of the glacial epoch; but when it finally passed away they too had disappeared, some having become altogether extinct while others continued to exist in more southern lands. Among the first class are the sabre-toothed tiger, the extinct Siberian camel (Merycotherium), three species of elephant, two of rhinoceros, two bears, five species of deer, and the gigantic beaver; among the latter are the hyæna, bear, and lion, which are considered to be only varieties of those which once inhabited Britain. Down to Pliocene times the flora of Europe was very similar to that which now prevails in Eastern Asia and Eastern North America. The late Professor Asa Gray has pointed out that hundreds of species of trees and shrubs of peculiar genera which still flourish in those countries are now completely wanting in Europe, and there is good reason to believe that these were exterminated during the glacial period, being cut off from a southern migration, first by the Alps, and then by the Mediterranean; whereas in eastern America and Asia the mountain chains run in a north and south direction, and there is nothing to prevent the flora from having been preserved by a southward migration into a milder region.44
Our next two chapters will be devoted to a discussion of the causes which brought about the glacial epoch, and that still more extraordinary climatic phenomenon—the mild climate and luxuriant vegetation of the Arctic zone. If my readers will follow me with the care and attention so difficult and interesting a problem requires and deserves, they will find that I have grappled with all the more important facts which have to be accounted for, and have offered what I believe is the first complete and sufficient explanation of them. The important influence of climatal changes on the dispersal of animals and plants is a sufficient justification for introducing such a discussion into the present volume.
CHAPTER VIII
THE CAUSES OF GLACIAL EPOCHS
Various Suggested Causes—Astronomical Causes of Changes of Climate—Difference of Temperature caused by Varying Distance of the Sun—Properties of Air and Water, Snow and Ice, in Relation to Climate—Effects of Snow on Climate—High Land and Great Moisture Essential to the Initiation of a Glacial Epoch—Perpetual Snow nowhere Exists on Lowlands—Conditions Determining the Presence or Absence of Perpetual Snow—Efficiency of Astronomical Causes in Producing Glaciation—Action of Meteorological causes in Intensifying Glaciation—Summary of Causes of Glaciation—Effect of Clouds and Fog in cutting off the Sun's Heat—South Temperate America as Illustrating the Influence of Astronomical Causes on Climate—Geographical Changes how far a Cause of Glaciation—Land acting as a Barrier to Ocean-currents—The theory of Interglacial Periods and their Probable Character—Probable Effect of Winter in Aphelion on the Climate of Britain—The Essential Principle of Climatal Change Restated—Probable Date of the last Glacial Epoch—Changes of the Sea-level dependent on Glaciation—The Planet Mars as bearing on the Theory of Excentricity as a Cause of Glacial Epochs.
No less than seven different causes have been at various times advanced to account for the glacial epoch and other changes of climate which the geological record proves to have taken place. These, as enumerated by Mr. Searles V. Wood, Jun., are as follows:—
1. A decrease in the original heat of our planet.
2. Changes in the obliquity of the ecliptic.
3. The combined effect of the precession of the equinoxes and of the excentricity of the earth's orbit.
4. Changes in the distribution of land and water.
5. Changes in the position of the earth's axis of rotation.
6. A variation in the amount of heat radiated by the sun.
7. A variation in the temperature of space.
Of the above, causes (1) and (2) are undoubted realities; but it is now generally admitted that they are utterly inadequate to produce the observed effects. Causes (5) (6) and (7) are all purely hypothetical, for though such changes may have occurred there is no evidence that they have occurred during geological time; and it is besides certain that they would not, either singly or combined, be adequate to explain the whole of the phenomena. There remain causes (3) and (4), which have the advantage of being demonstrated facts, and which are universally admitted to be capable of producing some effect of the nature required, the only question being whether, either alone or in combination, they are adequate to produce all the observed effects. It is therefore to these two causes that we shall confine our inquiry, taking first those astronomical causes whose complex and wide reaching effects have been so admirably explained and discussed by Dr. Croll in numerous papers and in his work—"Climate and Time in their Geological Relations."
DIAGRAM SHOWING THE ALTERED POSITION OF THE POLES AT INTERVALS OF 10,500 YEARS PRODUCED BY THE PRECESSION OF THE EQUINOXES AND THE MOTION OF THE APHELION; AND ITS EFFECT ON CLIMATE DURING A PERIOD OF HIGH EXCENTRICITY.
Astronomical Causes of Changes of Climate.—The earth moves in an elliptical orbit round the sun, which is situated in one of the foci of the ellipse, so that the distance of the sun from us varies during the year to a considerable amount. Strange to say we are now three millions of miles nearer to the sun in winter than in summer, while the reverse is the case in the southern hemisphere; and this must have some effect in making our northern winters less severe than those of the south temperate zone. But the earth moves more rapidly in that part of its orbit which is nearer to the sun, so that our winter is not only milder, but several days shorter, than that of the southern hemisphere. The distribution of land and sea and other local causes prevent us from making any accurate estimate of the effects due to these differences; but there can be no doubt that if our winter were as long as our summer is now and we were also three million miles further from the sun at the former period, a very decided difference of climate would result—our winter would be colder and longer, our summer hotter and shorter. Now there is a combination of astronomical revolutions (the precession of the equinoxes and the motion of the aphelion) which actually brings this change about every 10,500 years, so that after this interval the condition of the two hemispheres is reversed as regards nearness to the sun in summer, and comparative duration of summer and winter; and this change has been going on throughout
44
This subject is admirably discussed in Professor Asa Gray's Lecture on "Forest Geography and Archæology" in the